Thanks for your comments! I will keep up the work with this sprite and take your advice in account. I agree with you (pawige) about the colors, one or two of them had to be added late in the process since I had choose a bad palette from the beginning...
I think this could be improved a lot if you dropped most of the dithering. You could smooth everything out, especially the body, to get more solid forms. Also, you've got a couple colors that are rather close to each other in brightness, I would try to evenly space them out a bit more, you'd get a nicer overall look. I like the cracks and damage on the wings, the skulls are cool too, though they would also benefit from less filler dithering.
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