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Looks good, I like the non-color-reduced version better though. The environments are pretty cool, but the dragons look kind of wonky anatomy- and design-wise.
Not bad I think Guthrie could use some work though. Guthrie's tail looks really flat, and the paws and hind legs don't look right, although it might be just me. Other than what could I pinpoint, it's fine. I suggest you try to adjust the shading on the tail so it doesn't look flat. I can't find a good reference for tail shading though...
Also, the tileset reminds me of Earthbound. Was the Earthbound-esque look intentional?
Yeah, I was looking for a really nice, clean style. I had a hard time getting a style down at first so I referenced some Earthbound tiles and used that as a starting point.
Yeah, I'm aware my anatomy on Guthrie is a little wonky-looking. There's an in-universe explanation for that, with Guthrie being a stepping stone in the evolution of a pure dragon/bird hybrid. His descendants eventually loose their hind legs entirely but gain stronger front legs in the process. I'll give a shot on redoing the tail shading. Already have a few ideas for that.